by later hand, lower left in graphite: 21; by later hand, lower right in graphite: M. (erased); by later hand on mount, lower right in pen and black ink: An Animal by Parmegiano; by later hand on mount, lower right in graphite: 71; by later hand on mount, lower left in graphite: Colnaghi (faded); by unknown hand on mount, upper center verso in pen and brown ink: [TT?]; by unknown hand on mount, center verso in graphite: 4 / Parmeggiano; by unknown hand on mount, lower center in pencil: A5435
George John Spencer, 2nd Earl Spencer [1758-1834], Althorp (Lugt 1531)(his sale, Thomas Philipe, London, 14 June 1811, lot 530, as Parmigianino: "A camelion [sic]--red chalk, stumped--virgorously executed). Robert Clermont Witt [1872-1952], London (Lugt 2228b). P. & D. Colnaghi & Co., Ltd., London (inv. no. A5435 on verso of mount); Joseph F. McCrindle [1923-2008], New York; Joseph F. McCrindle Foundation, 2008; gift to NGA, 2009.
- Arcimboldo, 1526-1593: Nature and Fantasy, National Gallery of Art, Washington, D.C.; Palazzo Reale, Milan, 2010-2011, brochure no. 26 (shown only in Washington).
- The McCrindle Gift: A Distinguished Collection of Drawings and Watercolors. National Gallery of Art, Washington, 2012, 34-35, no. 6 (entry by Andaleeb Banta).